to LIAS
Botanische Staatssammlung München © 1995–2008
Data remain intellectual property of the data set authors.


  • Please read the Instructions for LIAS participants before entering or revising data.
  • Enter data only when button at the right side is green. When button is red, please wait for a couple of hours until database reorganization has been completed. [To verify the actual status of the pages, select the reload frame function in the context menu!]
  • You may use ColorRange for a numeric coding of colours.
  • These pages are XHTML compatible designed. May be that problems arise with older web browser versions.
  • Authorized LIAS contributors may check Personalized Deposits.

Go to: Data maintenance  –  Nomenclature: Taxonomic Hierarchy  –  Nomenclature: Type Information  –  Nomenclature: Taxonomic Literature  –  Biogeography  –  Ecology  –  Host or Phorophyte Taxonomy  –  Lichen Photobionts  –  Thallus Morphology: General Morphology  –  Thallus Morphology: Outline  –  Thallus Morphology: Upper Thallus Surface  –  Thallus Morphology: Lower Thallus Surface  –  Thallus Anatomy: Upper Cortex  –  Thallus Anatomy: Medulla  –  Reproduction  –  Ascocarp Morphology: General Ascocarp Morphology  –  Ascocarp Morphology: Wall  –  Ascocarp Morphology: Margin  –  Ascocarp Morphology: Disk  –  Ascocarp Anatomy: Exciple  –  Ascocarp Anatomy: General Amphithecium Anatomy  –  Ascocarp Anatomy: Amphithecial Cortex  –  Ascocarp Anatomy: Periphyses  –  Ascocarp Anatomy: Episamma  –  Ascocarp Anatomy: Epithecium  –  Ascocarp Anatomy: Hymenium  –  Ascocarp Anatomy: Interascal Hyphae  –  Ascocarp Anatomy: Subhymenium  –  Ascocarp Anatomy: Hypothecium  –  Ascocarp Anatomy: Asci  –  Ascocarp Anatomy: Ascospores  –  Conidioma Morphology  –  Conidioma Anatomy: Pycnidia  –  Conidioma Anatomy: Conidia  –  Chemistry: Secondary Metabolites  –  Chemistry: Spot Tests  –  Index of all items of the project


Taxon name:



Data maintenance   back to top


1. Data set source [mandatory] [mandatory character]     1. authentic     2. compiled     3. template     4. checklist item     5. standard item
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


2. Data set author(s) [mandatory] [mandatory character]     18. Egea J. M. & Torrente P.     60. Triebel D.  None selected () ()


3. Data set reviewer(s)     13. Egea J.M. & Torrente P.     58. Scholz P.     69. Triebel D.
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


4. Data set revision status [mandatory character]     2. revised  None selected () ()


5. Data set publication permission [mandatory character]     1. no     2. yes  None selected () ()


6. Data set publication date



Nomenclature: Taxonomic Hierarchy   back to top


7. Current taxonomic status     1. accepted     2. basionymous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


8. Taxonomic rank [mandatory] [mandatory character]     1. genus     2. species  None selected () ()


10. Taxon number

Mean: Upper range:


11. Genus name [mandatory] [mandatory character] () ()


12. Synonyms


13. Family name [mandatory] [mandatory character] () ()


14. Suborder name () ()


15. Order name () ()



Nomenclature: Type Information   back to top


16. Basionym


17. Type


19. Protologue


20. Type specimen label data



Nomenclature: Taxonomic Literature   back to top

21. Taxonomic notes


22. Taxonomic literature



Biogeography   back to top


27. Continent distribution [mandatory] [mandatory character]     1. Africa     2. Antarctica     3. Asia-Temperate     4. Australasia     5. Europe     6. Northern America     7. Southern America     8. Pacific     9. Asia-Tropical
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


38. Checklist records [mandatory] [mandatory character]     2. Austria     10. Great Britain     12. Iberic Peninsula     13. Italy     17. New Guinea     20. Sonoran Desert     26. United States and Canada (continental)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ecology   back to top


43. Life habit    ()  2. biotroph (inhabiting living organisms) ()


44. Lifestyle [mandatory] [mandatory character]     4. lichenized (= parasitic and / or mutual symbiotic with photobionts)     5. lichenicolous (= parasitic on lichens)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


46. Thallus-substrate interaction     1. endosubstratic     3. episubstratic
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


47. Habitat insolation     1. sun-exposed     2. partially shaded     3. fully shaded
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


48. Habitat moisture     1. dry     2. moist     3. wet
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Host or Phorophyte Taxonomy   back to top


71. Host fungus or lichen species



Lichen Photobionts   back to top


79. Primary photobiont presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


80. Primary photobiont kind     1. cyanobacterial     2. chlorococcal     3. trentepohlioid
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


86. Primary photobiont genus    ()  47. Trentepohlia ()


87. Primary photobiont family    ()  28. Trentepohliaceae ()


88. Primary photobiont order    ()  16. Trentepohliales ()


91. Secondary photobiont presence    ()  1. absent ()



Thallus Morphology: General Morphology   back to top


104. Thallus growth form [mandatory character]     1. indistinct     2. crustose
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


105. Thallus primary subdivision    ()  1. continuous ()


108. Crustose growth organization     1. leprose     2. granular     3. rimose     5. areolate (primarily areolate)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


133. Thallus parts thickness [categorical]     1. thin     2. thick
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


134. Thallus parts thickness [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm thick



Thallus Morphology: Outline   back to top


168. Primal thallus persistence     1. disappearing     2. persistent
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


169. Primal thallus development     1. vestigial     2. pronounced
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Morphology: Upper Thallus Surface   back to top


172. Upper surface colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     13. green     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     24. yellow     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     32. grey-brown (= greyish brown, brownish grey)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


176. Upper surface roughness     1. smooth     2. rough
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


177. Upper surface texture    ()  2. pulverulaceous ()


178. Upper surface special structures: presence     1. absent     2. present:
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


196. Upper surface pruina: presence     1. epruinose     2. pruinose
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


198. Upper surface pruina: colour     4. greenish     5. yellowish     7. reddish     8. whitish
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


201. Upper surface pseudocyphellae: presence    ()  1. absent ()


218. Upper surface cilia: presence    ()  1. eciliate ()


225. Upper surface hairs: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


230. Upper surface isidia: presence    ()  1. absent ()


252. Upper surface soralia: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


264. Upper surface soralia: differentiation     1. not confluent     2. confluent (but not fusing)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


275. Upper surface cephalodia: presence    ()  1. absent ()


278. Upper surface lobules: presence    ()  1. absent ()


279. Upper surface granules: presence    ()  1. absent ()


280. Upper surface thalloconidia: presence    ()  1. absent ()



Thallus Morphology: Lower Thallus Surface   back to top


286. Lower surface attachment kind    ()  1. by the whole lower surface ()


287. Lower surface special structures: presence     1. absent     2. present:
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


288. Lower surface pseudocyphellae: presence    ()  1. absent ()


295. Lower surface cyphellae: presence    ()  1. absent ()


300. Lower surface rhizines: presence    ()  1. absent ()


319. Lower surface thalloconidia: presence    ()  1. absent ()


325. Lower surface cavernae: presence    ()  1. absent ()


326. Lower surface tomentum: presence    ()  1. absent ()



Thallus Anatomy: Upper Cortex   back to top


329. Upper cortex: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


331. Upper cortex height [percentage]

Mean: Unit:
% of thallus width

332. Upper cortex height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high



Thallus Anatomy: Medulla   back to top


393. Medulla euamyloid reactivity     1. in Lugol's solution negative     2. in Lugol's solution positive
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


394. Medulla hemiamyloid reactivity     1. not hemiamyloid     2. hemiamyloid
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


395. Medulla iodine reaction intensity     1. weakly euamyloid     2. strongly euamyloid
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


396. Medulla iodine reaction colour     1. I + blue     2. I + violet     3. I + red     4. I + reddish yellow
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Reproduction   back to top


414. Ascomata presence [mandatory] [mandatory character]     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Morphology: General Ascocarp Morphology   back to top


418. Ascocarp structure [mandatory character]     1. apothecioid     2. perithecioid     3. cleistothecioid
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


420. Ascocarp outline     1. orbicular     3. irregular     4. linear     5. stellate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


421. Ascocarp location {on main thallus}    ()  2. laminal ()


426. Ascocarp host dependency     1. host-independent     2. host-dependent
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


427. Ascocarp location {on host}     5. within the ascocarp margin of the host     6. within the ascocarp disks of the host
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


428. Ascocarp emergence    ()  1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()


429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


430. Ascocarp basis     1. not constricted (flat below)     2. slightly constricted (moderately conical at the base)     3. strongly constricted (strongly conical at the base)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


431. Ascocarp abundance [categorical]     1. sparse     2. moderately abundant     3. abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


436. Ascocarp ascostroma formation     2. pseudostromatic     3. stromatic
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


438. {orbicular} ascocarp diameter [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm in diam.



Ascocarp Morphology: Wall   back to top


443. Ascocarp wall: carbonization     1. not carbonized     2. carbonized only in parts     3. carbonized around the ostiole, lacking a clypeus     4. with a carbonized black clypeus around the ostiole
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


444. Ascocarp wall: fusion {of neighboured perithecia}     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Morphology: Margin   back to top


448. Ascocarp margin: structure     4. crenulate     5. granulose
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


449. Ascocarp margin: roughness    ()  1. smooth ()


451. Ascocarp margin: persistency     1. persistent     2. excluded
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


452. Ascocarp margin: development     1. indistinct (thin)     2. distinct (moderately thickened)     3. prominent (thick)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


453. Ascocarp margin: colour     3. pale grey (= greyish white, whitish grey)     6. black     7. red     10. brownish red (= reddish brown)     13. green     24. yellow     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


457. Ascocarp margin: width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


458. Ascocarp margin: pruina abundance     1. epruinose     2. slightly pruinose     3. distinctly pruinose
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


462. Ascocarp margin: surface structures presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


468. Ascocarp margin: external filaments presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Morphology: Disk   back to top


487. Ascocarp disk: curvature     1. urceolate (strongly concave)     2. excavate (concave)     3. plane (expanded)     4. weakly convex     5. strongly convex
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


488. Ascocarp disk: colour     6. black     8. pale red (= reddish white, whitish red, incl. pink, rosé)     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


489. Ascocarp disk: exposure     1. not exposed     3. fully exposed
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


491. Ascocarp disk: pruina presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


492. Ascocarp disk: pruina abundance     1. scarce     2. abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


493. Ascocarp disk: pruina colour     1. white     2. yellow greenish     3. green     4. yellow     5. red     6. brownish red (= reddish brown)     7. pale grey (= whitish grey, greyish white)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Exciple   back to top


497. Ascocarp excipulum: plectenchyma structure    ()  2. paraplectenchymatous ()


498. Ascocarp excipulum: hyphal orientation     1. parallel (radiating outwards)     2. net-like (intricately interwoven, with no predominant orientation)     3. not easily discerned (strongly gelatinized or carbonized)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


504. Ascocarp excipulum: pigment carbonization     1. biatorine (not carbonized)     2. lecideine (carbonized)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


505. Ascocarp excipulum: layer pigmentation     2. white     4. grey     6. black     10. brownish red (= reddish brown)     13. green     16. brownish green (= greenish brown, incl. olive, olivaceous)     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: