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Botanische Staatssammlung München © 1995–2005
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Go to: Data maintenance  –  Nomenclature: Taxonomic Hierarchy  –  Nomenclature: Type Information  –  Nomenclature: Taxonomic Literature  –  Biogeography  –  Ecology  –  Host or Phorophyte Taxonomy  –  Lichen Photobionts  –  Thallus Morphology: General Morphology  –  Thallus Morphology: Outline  –  Thallus Morphology: Upper Thallus Surface  –  Thallus Anatomy: Upper Cortex  –  Thallus Anatomy: Medulla  –  Reproduction  –  Ascocarp Morphology: General Ascocarp Morphology  –  Ascocarp Morphology: Wall  –  Ascocarp Morphology: Margin  –  Ascocarp Morphology: Disk  –  Ascocarp Anatomy: Exciple  –  Ascocarp Anatomy: General Amphithecium Anatomy  –  Ascocarp Anatomy: Amphithecial Cortex  –  Ascocarp Anatomy: Periphyses  –  Ascocarp Anatomy: Episamma  –  Ascocarp Anatomy: Epithecium  –  Ascocarp Anatomy: Hymenium  –  Ascocarp Anatomy: Interascal Hyphae  –  Ascocarp Anatomy: Subhymenium  –  Ascocarp Anatomy: Hypothecium  –  Ascocarp Anatomy: Asci  –  Ascocarp Anatomy: Ascospores  –  Conidioma Morphology  –  Conidioma Anatomy: Pycnidia  –  Conidioma Anatomy: Conidia  –  Chemistry: Secondary Metabolites  –  Chemistry: Spot Tests  –  Index of all items of the project

Taxon name:


Data maintenance   back to top
1. Data set source [mandatory] [mandatory character]     1. authentic     2. compiled     3. template     5. standard item
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2. Data set author(s) [mandatory] [mandatory character]     18. Egea J. M. & Torrente P.  None selected () ()
5. Data set publication permission [mandatory character]     1. no     2. yes  None selected () ()
6. Data set publication date


Nomenclature: Taxonomic Hierarchy   back to top
7. Current taxonomic status     1. accepted     2. basionymous
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8. Taxonomic rank [mandatory] [mandatory character]     2. species  None selected () ()
11. Genus name [mandatory] [mandatory character] () ()
12. Synonyms

13. Family name [mandatory] [mandatory character]     1. genus of unknown placement (incertae sedis)     88. Opegraphaceae Stizenb. (1862)  None selected () ()
14. Suborder name     1. family of unknown placement (incertae sedis)  None selected () ()
15. Order name     2. Arthoniales  None selected () ()

Nomenclature: Type Information   back to top
17. Type [mandatory character]


19. Protologue


20. Type specimen label data



Nomenclature: Taxonomic Literature   back to top

21. Taxonomic notes


22. Taxonomic literature


Biogeography   back to top
27. Continent distribution [mandatory] [mandatory character]     1. Africa     2. Antarctica     3. Asia-Temperate     4. Australasia     5. Europe     6. Northern America     7. Southern America     8. Pacific     9. Asia-Tropical
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38. Checklist records [mandatory]     12. Iberic Peninsula     20. Sonoran Desert
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Ecology   back to top
43. Life habit    ()  2. biotroph (inhabiting living organisms) ()
44. Lifestyle [mandatory] [mandatory character]     4. lichenized (= parasitic and / or mutual symbiotic with photobionts)     5. lichenicolous (= parasitic on lichens)
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45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)
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46. Thallus-substrate interaction     1. endosubstratic     3. episubstratic
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47. Habitat insolation     1. sun-exposed     2. partially shaded     3. fully shaded
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48. Habitat moisture     1. dry     2. moist     3. wet
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53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
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Host or Phorophyte Taxonomy   back to top
71. Host fungus or lichen species


Lichen Photobionts   back to top
79. Primary photobiont presence     1. absent     2. present
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80. Primary photobiont kind     1. cyanobacterial     2. chlorococcal     3. trentepohlioid
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Thallus Morphology: General Morphology   back to top
104. Thallus growth form [mandatory character]     1. indistinct     2. crustose
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105. Thallus primary subdivision    ()  1. continuous ()
108. Crustose growth organization     1. leprose     2. granular     3. rimose     5. areolate (primarily areolate)
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133. Thallus parts thickness [categorical]     1. thin     2. thick
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134. Thallus parts thickness [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm thick

Thallus Morphology: Outline   back to top
168. Primal thallus persistence     1. disappearing     2. persistent
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169. Primal thallus development     1. vestigial     2. pronounced
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Thallus Morphology: Upper Thallus Surface   back to top
172. Upper surface colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     24. yellow     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     32. grey-brown (= greyish brown, brownish grey)
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176. Upper surface roughness     1. smooth     2. rough
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177. Upper surface texture    ()  2. pulverulaceous ()
178. Upper surface special structures: presence     1. absent     2. present:
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196. Upper surface pruina: presence     1. epruinose     2. pruinose
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198. Upper surface pruina: colour     4. greenish     5. yellowish     7. reddish     8. whitish
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225. Upper surface hairs: presence     1. absent     2. present
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252. Upper surface soralia: presence     1. absent     2. present
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264. Upper surface soralia: differentiation     1. not confluent     2. confluent (but not fusing)
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Thallus Anatomy: Upper Cortex   back to top
329. Upper cortex: presence     1. absent     2. present
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331. Upper cortex height [percentage]
Mean: Unit:
% of thallus width

332. Upper cortex height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high

Thallus Anatomy: Medulla   back to top
393. Medulla euamyloid reactivity     1. in Lugol's solution negative     2. in Lugol's solution positive
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394. Medulla hemiamyloid reactivity     1. not hemiamyloid     2. hemiamyloid
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395. Medulla iodine reaction intensity     1. weakly euamyloid     2. strongly euamyloid
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396. Medulla iodine reaction colour     1. I + blue     2. I + violet     3. I + red     4. I + reddish yellow
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Reproduction   back to top
414. Ascomata presence [mandatory] [mandatory character]     1. absent     2. present
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Ascocarp Morphology: General Ascocarp Morphology   back to top
418. Ascocarp structure [mandatory character]     1. apothecioid     2. perithecioid     3. cleistothecioid
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420. Ascocarp outline     1. orbicular     3. irregular     4. linear     5. stellate
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426. Ascocarp host dependency     1. host-independent     2. host-dependent
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427. Ascocarp location {on host}     5. within the ascocarp margin of the host     6. within the ascocarp disks of the host
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428. Ascocarp emergence    ()  1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()
429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile
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430. Ascocarp basis     1. not constricted (flat below)     2. slightly constricted (moderately conical at the base)     3. strongly constricted (strongly conical at the base)
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431. Ascocarp abundance [categorical]     1. sparse     2. moderately abundant     3. abundant
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436. Ascocarp ascostroma formation     2. pseudostromatic     3. stromatic
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438. {orbicular} ascocarp diameter [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm in diam.

Ascocarp Morphology: Wall   back to top
443. Ascocarp wall: carbonization     1. not carbonized     2. carbonized only in parts     3. carbonized around the ostiole, lacking a clypeus     4. with a carbonized black clypeus around the ostiole
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444. Ascocarp wall: fusion {of neighboured perithecia}     1. absent     2. present
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Ascocarp Morphology: Margin   back to top
448. Ascocarp margin: structure     4. crenulate     5. granulose
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449. Ascocarp margin: roughness    ()  1. smooth ()
451. Ascocarp margin: persistency     1. persistent     2. excluded
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452. Ascocarp margin: development     1. indistinct (thin)     2. distinct (moderately thickened)     3. prominent (thick)
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453. Ascocarp margin: colour     3. pale grey (= greyish white, whitish grey)     6. black     7. red     10. brownish red (= reddish brown)     13. green     24. yellow     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)
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457. Ascocarp margin: width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide
458. Ascocarp margin: pruina abundance     1. epruinose     2. slightly pruinose     3. distinctly pruinose
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462. Ascocarp margin: surface structures presence     1. absent     2. present
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468. Ascocarp margin: external filaments presence     1. absent     2. present
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Ascocarp Morphology: Disk   back to top
487. Ascocarp disk: curvature     1. urceolate (strongly concave)     2. excavate (concave)     3. plane (expanded)     4. weakly convex     5. strongly convex
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488. Ascocarp disk: colour     6. black     8. pale red (= reddish white, whitish red, incl. pink, rosé)     30. brown     31. pale brown (= brownish white, whitish brown)
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489. Ascocarp disk: exposure     1. not exposed     3. fully exposed
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491. Ascocarp disk: pruina presence     1. absent     2. present
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492. Ascocarp disk: pruina abundance     1. scarce     2. abundant
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493. Ascocarp disk: pruina colour     1. white     2. yellow greenish     3. green     4. yellow     5. red     6. brownish red (= reddish brown)     7. pale grey (= whitish grey, greyish white)
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Ascocarp Anatomy: Exciple   back to top
497. Ascocarp excipulum: plectenchyma structure    ()  2. paraplectenchymatous ()
498. Ascocarp excipulum: hyphal orientation     1. parallel (radiating outwards)     2. net-like (intricately interwoven, with no predominant orientation)     3. not easily discerned (strongly gelatinized or carbonized)
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504. Ascocarp excipulum: pigment carbonization     1. biatorine (not carbonized)     2. lecideine (carbonized)
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505. Ascocarp excipulum: layer pigmentation     2. white     4. grey     6. black     10. brownish red (= reddish brown)     13. green     16. brownish green (= greenish brown, incl. olive, olivaceous)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)
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506. Ascocarp excipulum: crystal inspersion     1. not inspersed with crystals     2. inspersed with crystals
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Ascocarp Anatomy: General Amphithecium Anatomy   back to top
559. Amphithecial photobiont abundance     1. absent     2. sparse (eulecanorine)     3. abundant
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Ascocarp Anatomy: Amphithecial Cortex   back to top
566. Amphithecial cortex presence     1. absent     2. present
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567. Amphithecial cortex development     1. indistinct     2. distinct
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Ascocarp Anatomy: Periphyses   back to top
613. Periphyses presence     1. absent     2. present
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Ascocarp Anatomy: Episamma   back to top
622. Episamma presence     1. absent     2. present
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626. Episamma solubility     1. dissolving in 10% KOH     2. not dissolving in 10% KOH
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Ascocarp Anatomy: Epithecium   back to top
627. Epithecium presence     1. absent     2. present
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629. Epithecium height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high
633. Epithecial hyphae apical cell: width [categorical]     1. not swollen     2. slightly swollen     3. distinctly swollen
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634. Epithecial hyphae apical cell: width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide
638. Epithecial hyphae apical cell: pigment location     1. pigment cap (within the cell wall)     3. diffuse (outside the cell wall)
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639. Epithecial hyphae apical cell pigmentation {in H2O}     1. hyaline (translucent)     10. brownish red (= reddish brown)     16. brownish green (= greenish brown, incl. olive, olivaceous)     30. brown     31. pale brown (= brownish white, whitish brown)     33. dark brown (= brownish black, blackish brown)
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Ascocarp Anatomy: Hymenium   back to top
650. Hymenium persistence    ()  2. mazaedioid (asci soon disintregating) ()
653. Hymenium height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high
654. Hymenium pigmentation {in H2O}     2. white     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     31. pale brown (= brownish white, whitish brown)
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667. Hymenium euamyloid reaction     1. Lugol’s negative     2. Lugol’s positive
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668. Hymenium hemiamyloid reaction     1. not hemiamyloid     2. hemiamyloid
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Ascocarp Anatomy: Interascal Hyphae   back to top
673. Interascal hyphae presence     1. absent     2. present
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676. Interascal hyphae ramification     1. scarcely branched     2. distinctly branched
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677. Interascal hyphae anastomoses     1. not or scarcely anastomosed     2. distinctly anastomosed
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681. Interascal hyphae cell width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide

Ascocarp Anatomy: Subhymenium   back to top
684. Subhymenium height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high
699. Subhymenium pigmentation {in H2O}     2. white     3. pale grey (= greyish white, whitish grey)     10. brownish red (= reddish brown)     30. brown     31. pale brown (= brownish white, whitish brown)
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Ascocarp Anatomy: Hypothecium   back to top
707. Hypothecium height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high
708. Hypothecium pigmentation {in H2O}     2. white     4. grey     6. black     10. brownish red (= reddish brown)     13. green     16. brownish green (= greenish brown, incl. olive, olivaceous)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)
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711. Hypothecium pigment pattern     1. homogenous     2. inhomogenous
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Ascocarp Anatomy: Asci   back to top
751. Ascus shape     1. globose     4. broadly clavate     5. clavate     6. narrowly clavate     8. cylindrical
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752. Ascus stipitation     1. absent     2. indistinct     3. distinct
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753. Ascus length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long

754. Ascus width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide
757. Ascus tholus thickness     1. not thickened     2. thickened
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758. Ascus tholus iodine reactivity     1. Lugol’s negative     2. Lugol’s positive
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759. Ascus tholus iodine reaction kind     1. hemiamyloid     2. euamyloid
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760. Ascus tholus iodine reaction colour     1. + red     2. + light blue     3. + dark blue
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761. Ascus tholus amyloidy pattern     1. with amyloid tube     2. with amyloid cap
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767. Ascus ocular chamber presence     1. absent     2. present
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768. Ascus ocular chamber development     1. indistinct     2. narrow     3. broad
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769. Ascus dehiscence type     1. prototunicate     2. lecanoralean     3. unitunicate     4. verrucarial     5. bitunicate
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780. Endoascus {hemiamyloid} reactivity     1. not hemiamyloid     2. inner layer hemiamyloid     3. all layers hemiamyloid
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Ascocarp Anatomy: Ascospores   back to top
784. Ascospore number [categorical]     2. c. 4 per ascus     3. c. 8 per ascus
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786. Ascospore shape     1. globose     4. ellipsoid     5. broadly ellipsoid     6. ovoid     10. oblong-obtuse     11. oblong     13. cylindrical     16. fusiform     17. filiform     23. clavate     28. semilunate     31. reniform (incl. fabiform)     32. curved     35. allantoid     36. bifusiform (incl. biclavate)     39. irregular
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