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Botanische Staatssammlung München © 1995–2008
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Go to: Data maintenance  –  Nomenclature: Taxonomic Hierarchy  –  Nomenclature: Type Information  –  Nomenclature: Taxonomic Literature  –  Biogeography  –  Ecology  –  Host or Phorophyte Taxonomy  –  Thallus Morphology: General Morphology  –  Thallus Morphology: Thallus Size and Differentiation  –  Thallus Morphology: Outline  –  Thallus Morphology: Upper Thallus Surface  –  Thallus Anatomy: Upper Cortex  –  Thallus Anatomy: Photobiont Layer  –  Thallus Anatomy: Medulla  –  Reproduction  –  Ascocarp Morphology: General Ascocarp Morphology  –  Ascocarp Morphology: Margin  –  Ascocarp Morphology: Disk  –  Ascocarp Anatomy: Episamma  –  Ascocarp Anatomy: Epithecium  –  Ascocarp Anatomy: Hymenium  –  Ascocarp Anatomy: Interascal Hyphae  –  Ascocarp Anatomy: Asci  –  Ascocarp Anatomy: Ascospores  –  Conidioma Morphology  –  Conidioma Anatomy: Pycnidia  –  Conidioma Anatomy: Conidia  –  Chemistry: Secondary Metabolites  –  Chemistry: Spot Tests  –  Index of all items of the project


Taxon name:



Data maintenance   back to top


1. Data set source [mandatory] [mandatory character]     1. authentic     2. compiled     3. template     5. standard item
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2. Data set author(s) [mandatory] [mandatory character]     6. Bowler P. A.     30. Kashiwadani H.     42. Nash T.H.  None selected () ()


3. Data set reviewer(s)    ()  59. Schramm R. & Nash T.H. ()


4. Data set revision status [mandatory character]     2. revised  None selected () ()


5. Data set publication permission [mandatory character]     1. no  None selected () ()



Nomenclature: Taxonomic Hierarchy   back to top


7. Current taxonomic status     1. accepted     2. basionymous
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8. Taxonomic rank [mandatory] [mandatory character]     2. species  None selected () ()


11. Genus name [mandatory] [mandatory character]     637. Ramalina  None selected () ()


12. Synonyms


13. Family name [mandatory] [mandatory character]     111. Ramalinaceae Ag. (1821)  None selected () ()


15. Order name     9. Lecanorales  None selected () ()



Nomenclature: Type Information   back to top


17. Type



Nomenclature: Taxonomic Literature   back to top

21. Taxonomic notes


22. Taxonomic literature



Biogeography   back to top


23. Hemisphere distribution    ()  1. Northern hemispheric ()


24. Floristic province {= zonal distribution} [mandatory] [mandatory character]     1. holarctic     2. arctic     3. subarctic     4. boreal     5. Mediterranean     6. subtropical     7. pantropical     8. neotropical     9. palaeotropic     10. subantarctic     11. antarctic     14. cosmopolitan
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25. Altitude [mandatory] [mandatory character]     1. coastal     2. colline     3. submontane     4. montane     5. upper montane     6. subalpine     7. alpine
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38. Checklist records [mandatory] [mandatory character]     15. Mexico     20. Sonoran Desert     26. United States and Canada (continental)
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Ecology   back to top


43. Life habit    ()  2. biotroph (inhabiting living organisms) ()


44. Lifestyle [mandatory] [mandatory character]    ()  5. lichenicolous (= parasitic on lichens) ()


45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)     5. epiphyllous (on living leaves)     6. saxicolous (on rock)
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53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
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Host or Phorophyte Taxonomy   back to top


71. Host fungus or lichen species



Thallus Morphology: General Morphology   back to top


104. Thallus growth form [mandatory character]     2. crustose     3. foliose     4. fruticose
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105. Thallus primary subdivision     1. continuous     2. divided
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106. Thallus secondary subdivision     1. entire     2. subdivided
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108. Crustose growth organization     2. granular     8. subfruticose (minutely fruticose, dactyliform)
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109. Crustose growth structure     1. bullate     2. subsquamulose
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112. Foliose growth organization    ()  1. lobed ()


113. Foliose lobe widening     1. widening     2. parallel
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116. Foliose lobe orientation     1. horizontal     2. ascending     3. imbricate
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117. Foliose lobe front    ()  1. angular ()


118. Foliose lobe curvature     1. concave     2. subconcave     3. plane     4. subconvex     5. convex
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119. Fruticose growth organization     1. erect     4. subpendulous     5. pendulous
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121. Fruticose branch structure     1. terete     2. subterete     3. applanate (moderately flattened)     4. compressed (strongly flattened)     5. alate     6. striate     7. ridged
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123. Fruticose branch segmentation     1. not segmented     2. segmented
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126. Thallus parts imbrication     1. not imbricate     2. imbricate
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129. Thallus parts length [categorical]     1. short     2. long
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130. Thallus parts length [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm long


131. Thallus parts width [categorical]     1. narrow     2. wide
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132. Thallus parts width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm wide


133. Thallus parts thickness [categorical]     1. thin     2. thick
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134. Thallus parts thickness [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm thick



Thallus Morphology: Thallus Size and Differentiation   back to top


151. Overall thallus diameter [categorical]     1. very small     2. small     3. medium-sized     4. large     5. very large
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153. Overall thallus length [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
cm long


154. Thallus joints main branch {basal} constriction     1. basally not constricted     2. basally constricted
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155. Thallus joints secondary branch {basal} constriction     1. basally not constricted     2. basally constricted
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157. Thallus apex width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm wide



Thallus Morphology: Outline   back to top


164. Thallus margin structure     1. entire     2. sulcate     3. dentate     4. laciniate     5. cucculate     6. undulate     7. isidiate     8. phyllidiate     9. crenulate     10. dissected     11. lobulate
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165. Thallus margin orientation     1. down-turned     2. levelled     3. up-turned
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Thallus Morphology: Upper Thallus Surface   back to top


171. Upper surface aspect     1. verrucose     2. verruculose
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172. Upper surface colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     8. pale red (= reddish white, whitish red, incl. pink, rosé)     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     20. pale blue (= whitish blue, bluish white)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     31. pale brown (= brownish white, whitish brown)     34. orange (= reddish yellow, yellowish red)
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173. Upper surface convolution    ()  1. convoluted ()


174. Upper surface curvature     1. plane     2. subconcave     3. concave     4. subconvex     5. convex
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175. Upper surface glossiness     1. matt (opaque, dull)     2. glossy (shiny)     3. vitreous
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176. Upper surface roughness     1. smooth     2. rough
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178. Upper surface special structures: presence     1. absent     2. present:
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179. Upper surface fissures: presence     1. absent     2. present
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180. Upper surface fissures: arrangement     1. single     2. reticulate
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183. Upper surface fissures: location     1. central     2. peripheral
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184. Upper surface fissures: orientation     1. longitudinal     2. transverse     3. without predominant orientation     4. both transverse and longitudinal
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185. Upper surface fissures: position     1. on tubercles     2. on wrinkles     3. on folds     4. within pseudocyphellae     5. between wrinkles
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186. Upper surface pits: presence     1. absent     2. present
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189. Upper surface folds: presence     1. absent     2. present
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190. Upper surface folds: orientation     1. without particular orientation     2. longitudinally orientated     3. transversally orientated
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193. Upper surface patches: presence     1. absent     2. present
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195. Upper surface patches: outline     1. punctiform     2. orbicular     3. linear
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196. Upper surface pruina: presence     1. epruinose     2. pruinose
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197. Upper surface pruina: abundance     1. scarce     2. abundant
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198. Upper surface pruina: colour     2. greyish     4. greenish     5. yellowish     8. whitish
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218. Upper surface cilia: presence     1. eciliate     2. ciliate
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220. Upper surface cilia: position     2. apical     6. marginal
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222. Upper surface cilia: length [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm long


224. Upper surface cilia: apical brightness [categorical]     1. bright tipped     2. evenly coloured     3. dark tipped
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230. Upper surface isidia: presence     1. absent     2. present
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234. Upper surface isidia: arrangement     1. isolated     2. clustered
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235. Upper surface isidia: shape     4. inflated     5. cylindrical     7. spathulate
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237. Upper surface isidia: ramification     1. not branched     2. branched
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243. Upper surface papillae: presence     1. absent     2. present
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244. Upper surface papillae: shape     1. minute (barely distinct)     2. subhemispherical     3. verrucose     4. cylindrical
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252. Upper surface soralia: presence     1. absent     2. present
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253. Upper surface soralia: diameter [categorical]     1. minute     2. broad
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254. Upper surface soralia: diameter [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm in diam.


255. Upper surface soralia: colour     3. pale grey (= greyish white, whitish grey)     8. pale red (= reddish white, whitish red, incl. pink, rosé)     14. pale green (= greenish white, whitish green)     16. brownish green (= greenish brown, incl. olive, olivaceous)     20. pale blue (= whitish blue, bluish white)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     31. pale brown (= brownish white, whitish brown)     34. orange (= reddish yellow, yellowish red)
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256. Upper surface soralia: location     1. central     2. laminal     3. subperipheral     4. peripheral
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257. Upper surface soralia: position     1. apical     2. subapical     3. lateral
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258. Upper surface soralia: origin     1. "ab initio" (literally: from the the beginning, i.e., directly forming on anywhere on thallus surface, breaking through the cortex)     2. papillar (forming at the tips of papillae, i.e., elongated, finger-like projections)     3. tubercular (eroding from the tip of tubercles, warts)     4. pustular (erupting from pustules)     5. nodular (developing on contorted, knot-like structures of branches of fruticose lichens)     6. rugular (originating on wrinkles)     7. on folds (forming on thallus folds)     8. fissural (forming from cracks or fissures)     9. undular (forming on thallus undulations)     10. along scars of fibrils
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259. Upper surface soralia: shape     1. semicapitate     2. capitate     3. hooded (helmet-shaped)     4. spathulate     6. maculiform
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260. Upper surface soralia: outline     1. erose (diffuse, unspecific)     2. punctiform     3. irregular ("effigurate")     4. linear
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261. Upper surface soralia: elevation     1. concave     2. plane     3. prominent
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262. Upper surface soralia: excavation    ()  2. fully excavated ()


265. Upper surface soralia: arrangement     1. in isolated lines     2. reticulate fused
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266. Upper surface soralia: orientation     1. longitudinal     2. transversal
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268. Upper surface soralia: development {from isidia}     1. not isidiate     2. isidiate
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271. Upper surface tubercles: presence     1. absent     2. present
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272. Upper surface tubercles: size [categorical]     1. very small     2. small     3. large
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Thallus Anatomy: Upper Cortex   back to top


329. Upper cortex: presence     1. absent     2. present
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330. Upper cortex type     1. phenocortical     2. eucortical
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332. Upper cortex height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm high


334. Upper cortex plectenchyma structure     1. prosoplectenchymatous     2. paraplectenchymatous
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335. Upper cortex paraplectenchyma type    ()  1. textura globularis ()


337. Upper cortex hyphal orientation    ()  2. anticlinal ()


338. Upper cortex layer layer differentiation     1. one     2. two
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339. Upper cortex hyphal cell structure    ()  1. chondroid ()


342. Upper cortex hyphal cell strands     1. conical hyphal cell strands penetrating the photobiont layer     2. irregular hyphal cell strands     3. without distinct strands of hyphae
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349. Upper cortex pigment location     1. in upper layer     2. in lower layer
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360. Upper cortex epicortex: presence     1. absent     2. present
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367. Upper cortex outer layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous     3. scleroplectenchymatous
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371. Upper cortex inner layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous     3. scleroplectenchymatous
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Thallus Anatomy: Photobiont Layer   back to top


374. Photobiont layer: height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm thick


376. Photobiont layer: compartmentalization     1. continuous     2. clustered     3. in vaulted chambers     4. scattered
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Thallus Anatomy: Medulla   back to top


380. Medulla height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm thick


384. Medulla density     1. compact     2. dense     3. loose     4. very loose, becoming hollow
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388. Medulla hyphal arrangement     1. forming several bundles     2. forming a compact mass
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389. Medulla hyphal orientation     1. predominantly along the lobe axis     2. without particular orientation in upper part, predominantly along the lobe axis in lower part     3. without particular orientation
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393. Medulla euamyloid reactivity     1. in Lugol's solution negative     2. in Lugol's solution positive
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395. Medulla iodine reaction intensity     1. weakly euamyloid     2. strongly euamyloid
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396. Medulla iodine reaction colour     1. I + blue     2. I + violet
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Reproduction   back to top


414. Ascomata presence [mandatory] [mandatory character]     1. absent     2. present
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Ascocarp Morphology: General Ascocarp Morphology   back to top


421. Ascocarp location {on main thallus}     1. central     2. laminal     3. subperipheral     4. peripheral
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424. Ascocarp position {on primary thallus parts}     1. central     2. laminal     3. subperipheral     4. peripheral     5. subterminal     6. terminal
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429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile     3. substipitate     4. stipitate
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430. Ascocarp basis     1. not constricted (flat below)     2. slightly constricted (moderately conical at the base)     3. strongly constricted (strongly conical at the base)
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431. Ascocarp abundance [categorical]     1. sparse     2. moderately abundant     3. abundant
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