to LIAS
Botanische Staatssammlung München © 1995–2008
Data remain intellectual property of the data set authors.


  • Please read the Instructions for LIAS participants before entering or revising data.
  • Enter data only when button at the right side is green. When button is red, please wait for a couple of hours until database reorganization has been completed. [To verify the actual status of the pages, select the reload frame function in the context menu!]
  • You may use ColorRange for a numeric coding of colours.
  • These pages are XHTML compatible designed. May be that problems arise with older web browser versions.
  • Authorized LIAS contributors may check Personalized Deposits.

Go to: Data maintenance  –  Nomenclature: Taxonomic Hierarchy  –  Nomenclature: Type Information  –  Nomenclature: Taxonomic Literature  –  Biogeography  –  Ecology  –  Host or Phorophyte Taxonomy  –  Thallus Morphology: General Morphology  –  Thallus Morphology: Thallus Size and Differentiation  –  Thallus Morphology: Outline  –  Thallus Morphology: Upper Thallus Surface  –  Thallus Morphology: Lower Thallus Surface  –  Thallus Anatomy: Upper Cortex  –  Thallus Anatomy: Photobiont Layer  –  Thallus Anatomy: Medulla  –  Thallus Anatomy: Lower Cortex  –  Reproduction  –  Ascocarp Morphology: General Ascocarp Morphology  –  Ascocarp Morphology: Margin  –  Ascocarp Morphology: Disk  –  Ascocarp Anatomy: Exciple  –  Ascocarp Anatomy: Parathecium  –  Ascocarp Anatomy: General Amphithecium Anatomy  –  Ascocarp Anatomy: Lower Amphithecium  –  Ascocarp Anatomy: Amphithecial Cortex  –  Ascocarp Anatomy: Episamma  –  Ascocarp Anatomy: Epithecium  –  Ascocarp Anatomy: Hymenium  –  Ascocarp Anatomy: Interascal Hyphae  –  Ascocarp Anatomy: Subhymenium  –  Ascocarp Anatomy: Hypothecium  –  Ascocarp Anatomy: Asci  –  Ascocarp Anatomy: Ascospores  –  Conidioma Morphology  –  Conidioma Anatomy: Conidiophores  –  Conidioma Anatomy: Conidia  –  Chemistry: Secondary Metabolites  –  Chemistry: Spot Tests  –  Chemistry: UV-Fluorescence  –  (Return to the index of all items of the project)


Taxon name:



Data maintenance   back to top


1. Data set source [mandatory] [mandatory character]     1. authentic     2. compiled     3. template     5. standard item
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


2. Data set author(s) [mandatory] [mandatory character] () ()


3. Data set reviewer(s)     23. Helms G.     40. Marbach B.     45. Nash T.H.     61. Sheard J.
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


5. Data set publication permission [mandatory character]     1. no     2. yes  None selected () ()


6. Data set publication date



Nomenclature: Taxonomic Hierarchy   back to top


7. Current taxonomic status     1. accepted     2. basionymous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


8. Taxonomic rank [mandatory] [mandatory character]     2. species     3. subspecies     4. variety     5. form  None selected () ()


9. Currently accepted name


11. Genus name [mandatory] [mandatory character] () ()


12. Synonyms


13. Family name [mandatory] [mandatory character]     99. Physciaceae Zahlbr. (1898)  None selected () ()


14. Suborder name     5. Lecanorineae  None selected () ()


15. Order name     9. Lecanorales  None selected () ()



Nomenclature: Type Information   back to top


16. Basionym


17. Type


19. Protologue


20. Type specimen label data



Nomenclature: Taxonomic Literature   back to top

21. Taxonomic notes


22. Taxonomic literature



Biogeography   back to top


23. Hemisphere distribution    ()  1. Northern hemispheric ()


24. Floristic province {= zonal distribution} [mandatory] [mandatory character]     1. holarctic     2. arctic     3. subarctic     4. boreal     5. Mediterranean     6. subtropical     7. pantropical     8. neotropical     9. palaeotropic     10. subantarctic     11. antarctic     14. cosmopolitan     15. temperate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


25. Altitude [mandatory] [mandatory character]     1. coastal     2. colline     3. submontane     4. montane     5. upper montane     6. subalpine     7. alpine
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


27. Continent distribution [mandatory] [mandatory character]     1. Africa     2. Antarctica     3. Asia-Temperate     4. Australasia     5. Europe     6. Northern America     7. Southern America     8. Pacific     9. Asia-Tropical
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


38. Checklist records [mandatory] [mandatory character]     1. Australia     2. Austria     7. Germany     10. Great Britain     11. Guianas     13. Italy     15. Mexico     17. New Guinea     18. New Zealand     20. Sonoran Desert     21. Sweden and Norway     22. Thailand     26. United States and Canada (continental)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ecology   back to top


43. Life habit    ()  2. biotroph (inhabiting living organisms) ()


44. Lifestyle [mandatory] [mandatory character]     4. lichenized (= parasitic and / or mutual symbiotic with photobionts)     5. lichenicolous (= parasitic on lichens)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)     6. saxicolous (on rock)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


46. Thallus-substrate interaction     1. endosubstratic     3. episubstratic
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


47. Habitat insolation     1. sun-exposed     2. partially shaded     3. fully shaded
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


48. Habitat moisture     1. dry     2. moist     3. wet
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Host or Phorophyte Taxonomy   back to top


72. Host fungus or lichen genus     6. Aspicilia     17. Dimelaena     24. Lecanora     55. Rhizocarpon
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Morphology: General Morphology   back to top


104. Thallus growth form [mandatory character]     1. indistinct     2. crustose     3. foliose     4. fruticose
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


105. Thallus primary subdivision     1. continuous     2. divided
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


107. Thallus ramification pattern     1. isotomic-dichotomous     4. subdichotomous     5. irregular
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


108. Crustose growth organization     1. leprose     2. granular     3. rimose     5. areolate (primarily areolate)     6. placodioid     7. squamulose     8. subfruticose (minutely fruticose, dactyliform)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


109. Crustose growth structure    ()  2. subsquamulose ()


110. Crustose growth aggregation    ()  2. scattered (dispersed) ()


111. Crustose growth continuity     1. continuous throughout     2. dispersed along the periphery, continuous in the centre     3. dispersed throughout and distinctly seperate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


112. Foliose growth organization    ()  1. lobed ()


113. Foliose lobe widening     1. widening     2. parallel
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


114. Foliose lobe outline     1. elongate     2. lunate     3. linear     4. flabellate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


116. Foliose lobe orientation     1. horizontal     2. ascending     3. imbricate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


117. Foliose lobe front     1. angular     2. rounded
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


118. Foliose lobe curvature     1. concave     2. subconcave     3. plane     4. subconvex     5. convex
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


119. Fruticose growth organization     1. erect     4. subpendulous     5. pendulous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


121. Fruticose branch structure     1. terete     2. subterete     3. applanate (moderately flattened)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


122. Fruticose branch flattening     1. bifacial     2. unifacial
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


126. Thallus parts imbrication     1. not imbricate     2. imbricate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


129. Thallus parts length [categorical]     1. short     2. long
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


131. Thallus parts width [categorical]     1. narrow     2. wide
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


132. Thallus parts width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm wide


133. Thallus parts thickness [categorical]     1. thin     2. thick
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


134. Thallus parts thickness [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm thick



Thallus Morphology: Thallus Size and Differentiation   back to top


151. Overall thallus diameter [categorical]     1. very small     2. small     3. medium-sized     4. large
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


152. Overall thallus diameter [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
cm in diameter


156. Thallus apex width [categorical]     1. not distinctly broader than average     2. distinctly broader than average
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


158. Thallus apex curvature     1. not curved     3. curved upwards     4. curved downwards
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


159. Thallus apex outline     1. rounded     2. truncate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Morphology: Outline   back to top


160. Thallus perimeter outline     1. irregular     2. orbicular
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


161. Thallus perimeter zonation     1. simple     2. zonate     3. lobed     5. fimbriate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


162. Thallus perimeter delimitation     1. indistinct     2. distinct
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


164. Thallus margin structure     3. dentate     4. laciniate     8. phyllidiate     11. lobulate     13. crenate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


165. Thallus margin orientation     1. down-turned     2. levelled     3. up-turned
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


166. Thallus margin brightness [categorical]     1. paled     2. concolourous     3. darkened
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


167. Primal thallus differentiation     1. peripheral (forming a prothallus)     2. extending (forming a hypothallus)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


168. Primal thallus persistence     1. disappearing     2. persistent
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


170. Primal thallus colour     1. black     2. grey     6. brown     8. white
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Morphology: Upper Thallus Surface   back to top


172. Upper surface colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     21. grey-blue (= bluish grey, greyish blue)     24. yellow     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


175. Upper surface glossiness     1. matt (opaque, dull)     2. glossy (shiny)     3. vitreous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


176. Upper surface roughness     1. smooth     2. rough
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


177. Upper surface texture     2. pulverulaceous     3. byssoid
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


191. Upper surface maculae: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


192. Upper surface maculae: outline     1. punctiform     2. orbicular     3. linear     4. irregular
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


196. Upper surface pruina: presence     1. epruinose     2. pruinose
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


197. Upper surface pruina: abundance     1. scarce     2. abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


198. Upper surface pruina: colour     2. greyish     4. greenish     8. whitish
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


199. Upper surface pruina: distribution     1. continuous     2. patchy
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


200. Upper surface pruina: location     1. central     3. peripheral
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


201. Upper surface pseudocyphellae: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


204. Upper surface pseudocyphellae: arrangement     1. isolated     2. reticulate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


208. Upper surface pseudocyphellae: location     1. central     2. laminal     3. peripheral
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


210. Upper surface pseudocyphellae: position     1. submarginal     2. marginal
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


218. Upper surface cilia: presence     1. eciliate     2. ciliate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


219. Upper surface cilia: kind     1. spinulous     2. slender
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


225. Upper surface hairs: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


227. Upper surface hairs: location     2. apical     3. laminal     4. interspersed     5. marginal
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


228. Upper surface hairs position    ()  1. on thallus lobes or branches ()


230. Upper surface isidia: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


231. Upper surface isidia: location    ()  2. laminal ()


232. Upper surface isidia: position {across the surface}     1. marginal     2. submarginal
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


234. Upper surface isidia: arrangement     1. isolated     2. clustered
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


235. Upper surface isidia: shape     1. globular     3. pustular     5. cylindrical     6. clavate     7. spathulate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


237. Upper surface isidia: ramification     1. not branched     2. branched
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


238. Upper surface isidia: ramification pattern     1. sparse     2. coralloid     3. moderate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


240. Upper surface isidiomorphs: presence    ()  2. present ()


252. Upper surface soralia: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


255. Upper surface soralia: colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     10. brownish red (= reddish brown)     14. pale green (= greenish white, whitish green)     16. brownish green (= greenish brown, incl. olive, olivaceous)     20. pale blue (= whitish blue, bluish white)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     30. brown     32. grey-brown (= greyish brown, brownish grey)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


256. Upper surface soralia: location     1. central     2. laminal     3. subperipheral     4. peripheral
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


257. Upper surface soralia: position     1. apical     2. subapical     3. lateral
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


258. Upper surface soralia: origin    ()  3. tubercular (eroding from the tip of tubercles, warts) ()


259. Upper surface soralia: shape     1. semicapitate     2. capitate     3. hooded (helmet-shaped)     4. spathulate     5. labriform     6. maculiform     7. ear-shaped
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


260. Upper surface soralia: outline     1. erose (diffuse, unspecific)     2. punctiform     3. irregular ("effigurate")     4. linear
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


261. Upper surface soralia: elevation    ()  3. prominent ()


262. Upper surface soralia: excavation     1. slightly excavated     2. fully excavated
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


264. Upper surface soralia: differentiation     1. not confluent     2. confluent (but not fusing)     3. fusing
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


268. Upper surface soralia: development {from isidia}    ()  2. isidiate ()


269. Upper surface soredia: diameter [categorical]     1. grainy (coarse)     2. farinose (fine)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


276. Upper surface blastidia: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


277. Upper surface schizidia: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


278. Upper surface lobules: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Morphology: Lower Thallus Surface   back to top


281. Lower surface presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


282. Lower surface overall colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     10. brownish red (= reddish brown)     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     21. grey-blue (= bluish grey, greyish blue)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


283. Lower surface central colour     2. white     4. grey     6. black     30. brown
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


284. Lower surface peripheral colour     1. black     2. grey     6. brown     8. white     28. pale brown (= brownish white, whitish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


285. Lower surface attachment extent     2. loosely attached     3. firmly attached
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


286. Lower surface attachment kind     1. by the whole lower surface     2. by holdfasts     3. lacking
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


296. Lower surface wrinkles: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


297. Lower surface wrinkles: location     1. central     2. peripheral
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


300. Lower surface rhizines: presence     1. absent     2. present (rhizinate)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


301. Lower surface rhizines: location     1. central     3. peripheral
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


302. Lower surface rhizines: abundance     1. sparse     2. moderately abundant     3. abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


307. Lower surface rhizines: colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


309. Lower surface rhizines: projections     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


310. Lower surface rhizines: ramification     1. not branched     2. branched
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


311. Lower surface rhizines: ramification pattern     1. irregular     2. dichotomous     3. squarrose     4. sparsely furcate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


316. Lower surface soralia: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


326. Lower surface tomentum: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Anatomy: Upper Cortex   back to top


329. Upper cortex: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


331. Upper cortex height [percentage]

Min: Lower range: Mean: Upper range: Max: Unit:
% of thallus width

332. Upper cortex height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high


349. Upper cortex pigment location     1. in upper layer     2. in lower layer
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


367. Upper cortex outer layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous     3. scleroplectenchymatous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


371. Upper cortex inner layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous     3. scleroplectenchymatous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Anatomy: Photobiont Layer   back to top


375. Photobiont layer: zonation     1. single-layered     2. multi-layered
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Anatomy: Medulla   back to top


380. Medulla height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm thick


383. Medulla colour     2. white     7. red     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     34. orange (= reddish yellow, yellowish red)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


384. Medulla density     1. compact     2. dense     3. loose     4. very loose, becoming hollow
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


388. Medulla hyphal arrangement     1. forming several bundles     2. forming a compact mass
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Thallus Anatomy: Lower Cortex   back to top


397. Lower cortex presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


400. Lower cortex height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm thick


401. Lower cortex plectenchyma structure     1. paraplectenchymatous     2. prosoplectenchymatous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


404. Lower cortex hyphal orientation     1. anticlinal     2. periclinal
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Reproduction   back to top


414. Ascomata presence [mandatory] [mandatory character]     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Morphology: General Ascocarp Morphology   back to top


418. Ascocarp structure [mandatory character]    ()  1. apothecioid ()


420. Ascocarp outline     1. orbicular     3. irregular
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


421. Ascocarp location {on main thallus}     1. central     2. laminal     3. subperipheral     4. peripheral
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


424. Ascocarp position {on primary thallus parts}     5. subterminal     6. terminal
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


428. Ascocarp emergence    ()  1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()


429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile     3. substipitate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


430. Ascocarp basis     1. not constricted (flat below)     2. slightly constricted (moderately conical at the base)     3. strongly constricted (strongly conical at the base)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


431. Ascocarp abundance [categorical]     1. sparse     2. moderately abundant     3. abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


432. Ascocarp abundance [numerical]

Min: Max: Unit:
per thallus


437. Ascocarp size [categorical]     1. small     2. large
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


438. {orbicular} ascocarp diameter [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
mm in diam.



Ascocarp Morphology: Margin   back to top


447. Ascocarp margin: type     1. lecideine/biatorine     2. lecanorine (incl. zeorine)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


448. Ascocarp margin: structure     2. lobate     3. lobulate     4. crenulate     5. granulose     6. scabrose     7. verrucose     8. verruculose
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


449. Ascocarp margin: roughness    ()  1. smooth ()


451. Ascocarp margin: persistency     1. persistent     2. excluded
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


452. Ascocarp margin: development     1. indistinct (thin)     2. distinct (moderately thickened)     3. prominent (thick)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


457. Ascocarp margin: width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


460. Ascocarp margin: parathecial crown presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


461. Ascocarp margin: parathecial crown brightness [categorical]     1. darker than surrounding margin     2. concolourous with surrounding margin     3. brighter than surrounding margin
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


463. Ascocarp margin: soredia     1. absent     2. present (sorediate)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


464. Ascocarp margin: isidia     1. absent     2. present (isidiate)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


466. Ascocarp margin: cilia     1. absent     2. present (ciliate)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


467. Ascocarp margin: hairs     1. absent     2. present (hirsute)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Morphology: Disk   back to top


487. Ascocarp disk: curvature     1. urceolate (strongly concave)     2. excavate (concave)     3. plane (expanded)     4. weakly convex
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


488. Ascocarp disk: colour     6. black     21. grey-blue (= bluish grey, greyish blue)     22. brownish blue (= bluish brown)     30. brown     32. grey-brown (= greyish brown, brownish grey)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


491. Ascocarp disk: pruina presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


492. Ascocarp disk: pruina abundance     1. scarce     2. abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


493. Ascocarp disk: pruina colour     1. white     2. yellow greenish     7. pale grey (= whitish grey, greyish white)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Exciple   back to top


504. Ascocarp excipulum: pigment carbonization    ()  2. lecideine (carbonized) ()


505. Ascocarp excipulum: layer pigmentation     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     5. dark gray (= greyish black, blackish grey)     6. black     7. red     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     11. bluish red (= reddish blue, incl. violet, purple)     12. dark red (= blackish red, reddish black)     13. green     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     19. blue     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     22. brownish blue (= bluish brown)     23. dark blue (= blackish blue, bluish black)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     30. brown     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


506. Ascocarp excipulum: crystal inspersion     1. not inspersed with crystals     2. inspersed with crystals
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Parathecium   back to top


507. Parathecium width [categorical]     1. narrow (indistinct, reduced)     2. broad (distinct)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


508. Parathecium width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


509. Parathecium structure     1. paraplectenchymatous     2. prosoplectenchymatous, strongly glutinized
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


510. Parathecium pigmentation     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     5. dark gray (= greyish black, blackish grey)     6. black     7. red     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     11. bluish red (= reddish blue, incl. violet, purple)     12. dark red (= blackish red, reddish black)     13. green     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     19. blue     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     22. brownish blue (= bluish brown)     23. dark blue (= blackish blue, bluish black)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     30. brown     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)     34. orange (= yellowish red, reddish yellow)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


514. Parathecium glutinization    ()  2. strongly glutinized ()


515. Parathecium crystal inspersion     1. not inspersed with crystals     2. inspersed with crystals
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


516. Parathecium crystal solubility     1. dissolving in 10% KOH     2. not dissolving in 10% KOH
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: General Amphithecium Anatomy   back to top


530. Amphithecium presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


538. Amphithecium height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm high

540. Amphithecium width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


541. Amphithecium inspersion: type     1. not inspers     3. inspersed with crystals
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


544. Amphithecium inspersion: crystal size    ()  1. small (< 5 µm in diameter) ()


545. Amphithecium inspersion: crystal abundance    ()  2. abundant ()


546. Amphithecium inspersion: crystal solubility     1. not dissolving in 10% KOH     2. dissolving in 10% KOH
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


559. Amphithecial photobiont abundance     1. absent     2. sparse (eulecanorine)     3. abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Lower Amphithecium   back to top


563. Lower amphithecium height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm high


564. Lower amphithecium plectenchyma structure     1. pseudoparenchymatous     2. prosoplectenchymatous
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


565. Lower amphithecium prosoplectenchyma hyphal orientation    ()  2. anticlinal ()



Ascocarp Anatomy: Amphithecial Cortex   back to top


566. Amphithecial cortex presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


572. Amphithecial cortex width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm wide

574. Amphithecial cortex basal width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide at the base


604. Amphithecial cortex: outer layer: pigmentation     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     5. dark gray (= greyish black, blackish grey)     6. black     7. red     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     11. bluish red (= reddish blue, incl. violet, purple)     12. dark red (= blackish red, reddish black)     13. green     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     19. blue     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     22. brownish blue (= bluish brown)     23. dark blue (= blackish blue, bluish black)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     30. brown     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


608. Amphithecial cortex: inner layer: pigmentation     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     5. dark gray (= greyish black, blackish grey)     6. black     7. red     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     11. bluish red (= reddish blue, incl. violet, purple)     12. dark red (= blackish red, reddish black)     13. green     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     19. blue     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     22. brownish blue (= bluish brown)     23. dark blue (= blackish blue, bluish black)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     30. brown     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Episamma   back to top


622. Episamma presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Epithecium   back to top


627. Epithecium presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


629. Epithecium height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm high


630. Epithecial hyphae apical cell: shape     1. simple     2. spathulate     3. subulate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


639. Epithecial hyphae apical cell pigmentation {in H2O}     10. brownish red (= reddish brown)     16. brownish green (= greenish brown, incl. olive, olivaceous)     22. brownish blue (= bluish brown)     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


641. Epithecial hyphae apical cell: pigment solubility {in 10% KOH}     1. insoluble in 10% KOH     2. soluble in 10% KOH
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


642. Epithecial hyphae apical cell: pigment reactivity    ()  2. 10% KOH + ()



Ascocarp Anatomy: Hymenium   back to top


653. Hymenium height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm high


654. Hymenium pigmentation {in H2O}     2. white     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     31. pale brown (= brownish white, whitish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


659. Hymenium gel presence     1. absent     2. present (interascal hyphae gelatinized)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


661. Hymenium inspersion: type     1. not inspers     2. inspersed with oil droplets     3. inspersed with crystals
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


667. Hymenium euamyloid reaction     1. Lugol’s negative     2. Lugol’s positive
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Interascal Hyphae   back to top


676. Interascal hyphae ramification     1. scarcely branched     2. distinctly branched
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


677. Interascal hyphae anastomoses     1. not or scarcely anastomosed     2. distinctly anastomosed
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Subhymenium   back to top


684. Subhymenium height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm high


699. Subhymenium pigmentation {in H2O}     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     5. dark gray (= greyish black, blackish grey)     6. black     7. red     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     11. bluish red (= reddish blue, incl. violet, purple)     12. dark red (= blackish red, reddish black)     13. green     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     19. blue     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     22. brownish blue (= bluish brown)     23. dark blue (= blackish blue, bluish black)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     30. brown     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Hypothecium   back to top


707. Hypothecium height [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm high


708. Hypothecium pigmentation {in H2O}     2. white     3. pale grey (= greyish white, whitish grey)     5. dark gray (= greyish black, blackish grey)     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     22. brownish blue (= bluish brown)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


716. Hypothecium prosoplectenchyma type     2. serioplectenchymatous     9. dense textura intricata
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


717. Hypothecium hyphal orientation     2. anticlinal     3. irregular
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


721. Hypothecium inspersion type     1. not inspers     2. inspersed with oil droplets
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Ascocarp Anatomy: Asci   back to top


751. Ascus shape     4. broadly clavate     5. clavate     6. narrowly clavate     7. obclavate     8. cylindrical
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


753. Ascus length [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm long

754. Ascus width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


769. Ascus dehiscence type    ()  2. lecanoralean ()



Ascocarp Anatomy: Ascospores   back to top


782. Ascospore type     1. Beltraminia-type     2. Bicincta-type     3. Bischoffii-type     5. Callispora-type     7. Conradii-type     8. Diplotomma-type     9. Dirinaria-type     14. Milvina-type     15. Mischoblastia-type     16. Orcularia-type     17. Pachysporaria-type     19. Physcia-type     20. Physcia-type and Pachysporaria-type     21. Physconia-type     23. Rinodinella-type     27. Tunicata-type
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


784. Ascospore number [categorical]     1. 1–2 per ascus     2. c. 4 per ascus     3. c. 8 per ascus     4. 12-16 per ascus     5. 16-32 per ascus     6. more than 32 per ascus
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


785. Ascospore number [numerical]

Min: Mean: Upper range: Max: Unit:
per ascus


786. Ascospore shape     3. oval     4. ellipsoid     5. broadly ellipsoid     16. fusiform
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


787. Ascospore length [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm long

788. Ascospore width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


792. Ascospore septum presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


793. Ascospore septum orientation     1. transversely septate     2. muriform
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


795. Ascospore {median} septum number [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
-transversally septate


799. Ascospore septum formation     1. early septum formation (Physciaceae-type A)     2. late septum formation (Physciaceae-type B)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


800. Ascospore cell lumen size [categorical]     1. even (all cells of equal size)     2. constricted in the centre (apically and basally enlarged)     3. tapered (swollen in the centre)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


802. Ascospore wall width [categorical]     1. thin     2. thick
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


803. Ascospore wall width [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


804. Ascospore wall differentiation {visible in the light microscope}    ()  2. distinctly differentiated (into primary and secondary wall) ()


805. {apical} ascospore wall thickening     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


806. {lateral} ascospore wall thickening     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


807. {septal} ascospore wall thickening     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


809. Ascospore wall pigment development     1. remaining hyaline     2. becoming pigmented
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


810. Ascospore wall pigmentation     2. grey     4. pale brown (= brownish white, whitish brown)     5. middle brown     7. dark brown (= brownish black, blackish brown)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


812. Ascospore perispore ornamentation     1. not ornamented (psilate, smooth)     2. ornamented
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


813. Ascospore perispore surface structure     1. echinate     2. foveate     3. rugulate     4. striate     5. porate
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


814. Ascospore mucilaginous sheath width [categorical]     1. indistinctly halonate (mucilaginous sheath absent or thin)     2. distinctly halonate (with a thick mucilaginous sheath)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Conidioma Morphology   back to top


816. Conidiomata presence [mandatory character]     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


818. Conidiomata abundance     1. sparse     2. moderately abundant     3. strongly abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Conidioma Anatomy: Conidiophores   back to top


857. Conidiophore type {sensu Vobis, Braun, etc.}     7. Vobis-type I     8. Vobis-type II     9. Vobis-type III     10. Vobis-type IV     11. Vobis-type V     12. Vobis-type VI     13. Vobis-type VII     14. Vobis-type VIII
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Conidioma Anatomy: Conidia   back to top


873. Conidium shape     1. globose     2. subglobose     3. oval     4. ellipsoid     5. broadly ellipsoid     11. bacilliform     13. cylindrical     16. fusiform
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


879. Conidium length [numerical]

Min: Lower range: Mean: Upper range: Max: Unit:
µm long

880. Conidium width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide



Chemistry: Secondary Metabolites   back to top


913. Known secondary metabolites: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


914. Known secondary metabolites: location     2. only in the upper part of the cortex     4. throughout the thallus medulla     11. in soredia (or soralia)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


915. Known secondary metabolites: name     231. atranorin     250. chloroatranorin     336. gyrophoric acid     347. hopane-6a,22-diol     390. leucotylin     392. lichexanthone     442. miriquidic acid     470. norstictic acid     486. paludosic acid     491. pannarin     536. salazinic acid     537. scabrosin 4,4'-diacetate     538. scabrosin 4,4'-dibutyrate     539. scabrosin 4-acetate-4'-butyrate     540. scabrosin 4-acetate-4'-hexanoate     543. secalonic acid a     549. skyrin     559. stictic acid     588. variolaric acid
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


916. Known secondary metabolites: substance class     2. (higher) aliphatic acids     3. (anthra-)quinones     13. secalonic acids
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


917. Unknown secondary metabolites: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


919. Unknown secondary metabolites: provisional name    ()  1. 'Heterodermia unknown spathulin' ()


921. Calcium oxalate crystals: presence     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Chemistry: Spot Tests   back to top


926. Spot test reactions     1. absent     2. present
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


927. Upper surface K reactivity     1. – (negative)     2. + deep yellow     5. + red
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


928. Upper surface C reactivity     1. –     5. + red
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


930. Upper surface KC reactivity     1. –     5. + red
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


931. Upper surface PD reactivity     1. –     2. + yellow     8. + orange
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


932. Cortex K reactivity     1. –     2. + deep yellow     3. + bright yellow (citrine, lemon)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


935. Cortex KC reactivity     1. –     2. + deep yellow     3. + bright yellow (citrine, lemon)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


936. Cortex PD reactivity     1. –     2. + yellow
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


937. Medulla K reactivity     1. –     2. + yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     8. + orange
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


938. Medulla C reactivity     1. –     5. + red
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


940. Medulla KC reactivity     1. –     2. + yellow     3. + bright yellow (citrine, lemon)     5. + red     9. + orange
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


941. Medulla PD reactivity     1. –     2. + deep yellow     5. + red
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


942. Lower surface K reactivity     1. –     2. + deep yellow     5. + red
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


947. Soredia K reactivity     1. –     2. + deep yellow     3. + pink     4. + orange
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


949. Soredia KC reactivity     1. –     2. + deep yellow     3. + pink     4. + orange
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )



Chemistry: UV-Fluorescence   back to top


983. UV-fluorescence: upper surface     1. – (negative)     2. + (positive)
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )


 

To send data, please enter your user password or your full email address (required!):

 

 


(Return to the index of all items of the project)

In case that additional characters and states are required to be included in this data set, consult the LIAS Instructions to Participants and follow the procedures described there.

LIAS Home