Taxon name:

Data maintenance  

1. Data set source [mandatory] [mandatory character]     1. authentic     2. compiled     3. template     5. standard item
(Select a character state: 1 2 3 5 to add this modifier: )
(Select a character state: 1 2 3 5 to add this note/comment: )

2. Data set author(s) [mandatory] [mandatory character] () Bungartz F. Esslinger T. Helms G. Marbach B. Moberg R. Nash T.H. Schramm R. & Nash T. H. Sheard J. ()

3. Data set reviewer(s)     23. Helms G.     40. Marbach B.     45. Nash T.H.     61. Sheard J.
(Select a character state: 23 40 45 61 to add this modifier: )
(Select a character state: 23 40 45 61 to add this note/comment: )

5. Data set publication permission [mandatory character]     1. no     2. yes  None selected () ()

6. Data set publication date

Nomenclature: Taxonomic Hierarchy  

7. Current taxonomic status     1. accepted     2. basionymous
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

8. Taxonomic rank [mandatory] [mandatory character]     2. species     3. subspecies     4. variety     5. form  None selected () ()

9. Currently accepted name

11. Genus name [mandatory] [mandatory character] () Amandinea Anaptychia Australiaena Buellia Culbersonia Dermatiscum Dermiscellum Dimelaena Diploicia Diplotomma Dirinaria Hafellia Heterodermia Hyperphyscia Phaeophyscia Phaeorrhiza Physcia Physconia Pyxine Redonia Rinodina Rinodinella Santessonia Tornabea ()

12. Synonyms

13. Family name [mandatory] [mandatory character]     99. Physciaceae Zahlbr. (1898)  None selected () ()

14. Suborder name     5. Lecanorineae  None selected () ()

15. Order name     9. Lecanorales  None selected () ()

Nomenclature: Type Information  

16. Basionym


17. Type


19. Protologue


20. Type specimen label data



Nomenclature: Taxonomic Literature  

21. Taxonomic notes


22. Taxonomic literature

Biogeography  

23. Hemisphere distribution    ()  1. Northern hemispheric ()

24. Floristic province {= zonal distribution} [mandatory] [mandatory character]     1. holarctic     2. arctic     3. subarctic     4. boreal     5. Mediterranean     6. subtropical     7. pantropical     8. neotropical     9. palaeotropic     10. subantarctic     11. antarctic     14. cosmopolitan     15. temperate
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(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 14 15 to add this note/comment: )

25. Altitude [mandatory] [mandatory character]     1. coastal     2. colline     3. submontane     4. montane     5. upper montane     6. subalpine     7. alpine
(Select a character state: 1 2 3 4 5 6 7 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 to add this note/comment: )

27. Continent distribution [mandatory] [mandatory character]     1. Africa     2. Antarctica     3. Asia-Temperate     4. Australasia     5. Europe     6. Northern America     7. Southern America     8. Pacific     9. Asia-Tropical
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this note/comment: )

38. Checklist records [mandatory] [mandatory character]     1. Australia     2. Austria     7. Germany     10. Great Britain     11. Guianas     13. Italy     15. Mexico     17. New Guinea     18. New Zealand     20. Sonoran Desert     21. Sweden and Norway     22. Thailand     26. United States and Canada (continental)
(Select a character state: 1 2 7 10 11 13 15 17 18 20 21 22 26 to add this modifier: )
(Select a character state: 1 2 7 10 11 13 15 17 18 20 21 22 26 to add this note/comment: )

Ecology  

43. Life habit    ()  2. biotroph (inhabiting living organisms) ()

44. Lifestyle [mandatory] [mandatory character]     4. lichenized (= parasitic and / or mutual symbiotic with photobionts)     5. lichenicolous (= parasitic on lichens)
(Select a character state: 4 5 to add this modifier: )
(Select a character state: 4 5 to add this note/comment: )

45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)     6. saxicolous (on rock)
(Select a character state: 1 2 3 4 6 to add this modifier: )
(Select a character state: 1 2 3 4 6 to add this note/comment: )

46. Thallus-substrate interaction     1. endosubstratic     3. episubstratic
(Select a character state: 1 3 to add this modifier: )
(Select a character state: 1 3 to add this note/comment: )

47. Habitat insolation     1. sun-exposed     2. partially shaded     3. fully shaded
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

48. Habitat moisture     1. dry     2. moist     3. wet
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
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(Select a character state: 1 2 to add this note/comment: )

Host or Phorophyte Taxonomy  

72. Host fungus or lichen genus     6. Aspicilia     17. Dimelaena     24. Lecanora     55. Rhizocarpon
(Select a character state: 6 17 24 55 to add this modifier: )
(Select a character state: 6 17 24 55 to add this note/comment: )

Thallus Morphology: General Morphology  

104. Thallus growth form [mandatory character]     1. indistinct     2. crustose     3. foliose     4. fruticose
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(Select a character state: 1 2 3 4 to add this note/comment: )

105. Thallus primary subdivision     1. continuous     2. divided
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

107. Thallus ramification pattern     1. isotomic-dichotomous     4. subdichotomous     5. irregular
(Select a character state: 1 4 5 to add this modifier: )
(Select a character state: 1 4 5 to add this note/comment: )

108. Crustose growth organization     1. leprose     2. granular     3. rimose     5. areolate (primarily areolate)     6. placodioid     7. squamulose     8. subfruticose (minutely fruticose, dactyliform)
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(Select a character state: 1 2 3 5 6 7 8 to add this note/comment: )

109. Crustose growth structure    ()  2. subsquamulose ()

110. Crustose growth aggregation    ()  2. scattered (dispersed) ()

111. Crustose growth continuity     1. continuous throughout     2. dispersed along the periphery, continuous in the centre     3. dispersed throughout and distinctly seperate
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

112. Foliose growth organization    ()  1. lobed ()

113. Foliose lobe widening     1. widening     2. parallel
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

114. Foliose lobe outline     1. elongate     2. lunate     3. linear     4. flabellate
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

116. Foliose lobe orientation     1. horizontal     2. ascending     3. imbricate
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

117. Foliose lobe front     1. angular     2. rounded
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

118. Foliose lobe curvature     1. concave     2. subconcave     3. plane     4. subconvex     5. convex
(Select a character state: 1 2 3 4 5 to add this modifier: )
(Select a character state: 1 2 3 4 5 to add this note/comment: )

119. Fruticose growth organization     1. erect     4. subpendulous     5. pendulous
(Select a character state: 1 4 5 to add this modifier: )
(Select a character state: 1 4 5 to add this note/comment: )

121. Fruticose branch structure     1. terete     2. subterete     3. applanate (moderately flattened)
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

122. Fruticose branch flattening     1. bifacial     2. unifacial
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

126. Thallus parts imbrication     1. not imbricate     2. imbricate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

129. Thallus parts length [categorical]     1. short     2. long
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

131. Thallus parts width [categorical]     1. narrow     2. wide
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

132. Thallus parts width [numerical]

133. Thallus parts thickness [categorical]     1. thin     2. thick
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

134. Thallus parts thickness [numerical]

Thallus Morphology: Thallus Size and Differentiation  

151. Overall thallus diameter [categorical]     1. very small     2. small     3. medium-sized     4. large
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

152. Overall thallus diameter [numerical]

156. Thallus apex width [categorical]     1. not distinctly broader than average     2. distinctly broader than average
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

158. Thallus apex curvature     1. not curved     3. curved upwards     4. curved downwards
(Select a character state: 1 3 4 U to add this modifier: )
(Select a character state: 1 3 4 U to add this note/comment: )

159. Thallus apex outline     1. rounded     2. truncate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Thallus Morphology: Outline  

160. Thallus perimeter outline     1. irregular     2. orbicular
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

161. Thallus perimeter zonation     1. simple     2. zonate     3. lobed     5. fimbriate
(Select a character state: 1 2 3 5 to add this modifier: )
(Select a character state: 1 2 3 5 to add this note/comment: )

162. Thallus perimeter delimitation     1. indistinct     2. distinct
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

164. Thallus margin structure     3. dentate     4. laciniate     8. phyllidiate     11. lobulate     13. crenate
(Select a character state: 3 4 8 11 13 to add this modifier: )
(Select a character state: 3 4 8 11 13 to add this note/comment: )

165. Thallus margin orientation     1. down-turned     2. levelled     3. up-turned
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

166. Thallus margin brightness [categorical]     1. paled     2. concolourous     3. darkened
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

167. Primal thallus differentiation     1. peripheral (forming a prothallus)     2. extending (forming a hypothallus)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

168. Primal thallus persistence     1. disappearing     2. persistent
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

170. Primal thallus colour     1. black     2. grey     6. brown     8. white
(Select a character state: 1 2 6 8 to add this modifier: )
(Select a character state: 1 2 6 8 to add this note/comment: )

Thallus Morphology: Upper Thallus Surface  

172. Upper surface colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     10. brownish red (= reddish brown)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     21. grey-blue (= bluish grey, greyish blue)     24. yellow     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)
(Select a character state: 2 3 4 6 8 9 10 15 16 21 24 26 27 30 31 32 33 to add this modifier: )
(Select a character state: 2 3 4 6 8 9 10 15 16 21 24 26 27 30 31 32 33 to add this note/comment: )

175. Upper surface glossiness     1. matt (opaque, dull)     2. glossy (shiny)     3. vitreous
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

176. Upper surface roughness     1. smooth     2. rough
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

177. Upper surface texture     2. pulverulaceous     3. byssoid
(Select a character state: 2 3 to add this modifier: )
(Select a character state: 2 3 to add this note/comment: )

191. Upper surface maculae: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

192. Upper surface maculae: outline     1. punctiform     2. orbicular     3. linear     4. irregular
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

196. Upper surface pruina: presence     1. epruinose     2. pruinose
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

197. Upper surface pruina: abundance     1. scarce     2. abundant
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

198. Upper surface pruina: colour     2. greyish     4. greenish     8. whitish
(Select a character state: 2 4 8 to add this modifier: )
(Select a character state: 2 4 8 to add this note/comment: )

199. Upper surface pruina: distribution     1. continuous     2. patchy
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

200. Upper surface pruina: location     1. central     3. peripheral
(Select a character state: 1 3 to add this modifier: )
(Select a character state: 1 3 to add this note/comment: )

201. Upper surface pseudocyphellae: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

204. Upper surface pseudocyphellae: arrangement     1. isolated     2. reticulate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

208. Upper surface pseudocyphellae: location     1. central     2. laminal     3. peripheral
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

210. Upper surface pseudocyphellae: position     1. submarginal     2. marginal
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

218. Upper surface cilia: presence     1. eciliate     2. ciliate
(Select a character state: 1 2 U to add this modifier: )
(Select a character state: 1 2 U to add this note/comment: )

219. Upper surface cilia: kind     1. spinulous     2. slender
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

225. Upper surface hairs: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

227. Upper surface hairs: location     2. apical     3. laminal     4. interspersed     5. marginal
(Select a character state: 2 3 4 5 to add this modifier: )
(Select a character state: 2 3 4 5 to add this note/comment: )

228. Upper surface hairs position    ()  1. on thallus lobes or branches ()

230. Upper surface isidia: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

231. Upper surface isidia: location    ()  2. laminal ()

232. Upper surface isidia: position {across the surface}     1. marginal     2. submarginal
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

234. Upper surface isidia: arrangement     1. isolated     2. clustered
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

235. Upper surface isidia: shape     1. globular     3. pustular     5. cylindrical     6. clavate     7. spathulate
(Select a character state: 1 3 5 6 7 to add this modifier: )
(Select a character state: 1 3 5 6 7 to add this note/comment: )

237. Upper surface isidia: ramification     1. not branched     2. branched
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

238. Upper surface isidia: ramification pattern     1. sparse     2. coralloid     3. moderate
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

240. Upper surface isidiomorphs: presence    ()  2. present ()

252. Upper surface soralia: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

255. Upper surface soralia: colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     10. brownish red (= reddish brown)     14. pale green (= greenish white, whitish green)     16. brownish green (= greenish brown, incl. olive, olivaceous)     20. pale blue (= whitish blue, bluish white)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     30. brown     32. grey-brown (= greyish brown, brownish grey)
(Select a character state: 2 3 4 10 14 16 20 24 25 26 27 30 32 to add this modifier: )
(Select a character state: 2 3 4 10 14 16 20 24 25 26 27 30 32 to add this note/comment: )

256. Upper surface soralia: location     1. central     2. laminal     3. subperipheral     4. peripheral
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

257. Upper surface soralia: position     1. apical     2. subapical     3. lateral
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

258. Upper surface soralia: origin    ()  3. tubercular (eroding from the tip of tubercles, warts) ()

259. Upper surface soralia: shape     1. semicapitate     2. capitate     3. hooded (helmet-shaped)     4. spathulate     5. labriform     6. maculiform     7. ear-shaped
(Select a character state: 1 2 3 4 5 6 7 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 to add this note/comment: )

260. Upper surface soralia: outline     1. erose (diffuse, unspecific)     2. punctiform     3. irregular ("effigurate")     4. linear
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

261. Upper surface soralia: elevation    ()  3. prominent ()

262. Upper surface soralia: excavation     1. slightly excavated     2. fully excavated
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

264. Upper surface soralia: differentiation     1. not confluent     2. confluent (but not fusing)     3. fusing
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

268. Upper surface soralia: development {from isidia}    ()  2. isidiate ()

269. Upper surface soredia: diameter [categorical]     1. grainy (coarse)     2. farinose (fine)
(Select a character state: 1 2 U to add this modifier: )
(Select a character state: 1 2 U to add this note/comment: )

276. Upper surface blastidia: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

277. Upper surface schizidia: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

278. Upper surface lobules: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Thallus Morphology: Lower Thallus Surface  

281. Lower surface presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

282. Lower surface overall colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     10. brownish red (= reddish brown)     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     21. grey-blue (= bluish grey, greyish blue)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: 2 3 4 6 10 14 15 21 24 25 26 27 30 31 to add this modifier: )
(Select a character state: 2 3 4 6 10 14 15 21 24 25 26 27 30 31 to add this note/comment: )

283. Lower surface central colour     2. white     4. grey     6. black     30. brown
(Select a character state: 2 4 6 30 to add this modifier: )
(Select a character state: 2 4 6 30 to add this note/comment: )

284. Lower surface peripheral colour     1. black     2. grey     6. brown     8. white     28. pale brown (= brownish white, whitish brown)
(Select a character state: 1 2 6 8 28 to add this modifier: )
(Select a character state: 1 2 6 8 28 to add this note/comment: )

285. Lower surface attachment extent     2. loosely attached     3. firmly attached
(Select a character state: 2 3 to add this modifier: )
(Select a character state: 2 3 to add this note/comment: )

286. Lower surface attachment kind     1. by the whole lower surface     2. by holdfasts     3. lacking
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

296. Lower surface wrinkles: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

297. Lower surface wrinkles: location     1. central     2. peripheral
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

300. Lower surface rhizines: presence     1. absent     2. present (rhizinate)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

301. Lower surface rhizines: location     1. central     3. peripheral
(Select a character state: 1 3 to add this modifier: )
(Select a character state: 1 3 to add this note/comment: )

302. Lower surface rhizines: abundance     1. sparse     2. moderately abundant     3. abundant
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

307. Lower surface rhizines: colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     30. brown     31. pale brown (= brownish white, whitish brown)
(Select a character state: 2 3 4 6 30 31 to add this modifier: )
(Select a character state: 2 3 4 6 30 31 to add this note/comment: )

309. Lower surface rhizines: projections     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

310. Lower surface rhizines: ramification